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参考解析

解析:
based。考点:动词。空格后的句子为定语从句,用来修饰“languages”,be based on“根据”,这里省略be,补充完整为which are based on…。这句话意为“其他的研究者也教过猩猩使用复杂的“语言”,这些语言是基于彩色的图形或电脑键盘”。

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